Hackelia deflexa (2000)
BORAGINACEAE
Hackelia deflexa (Wahlenb.) Opiz
U.S.A., South Dakota, Custer County: Black Hills: Black Elk Wilderness: divide between Iron Creek Trail and Grizzly Bear Creek Trail near junction with Harney Peak Trail, ca 1.4 air mi SSE of Harney Peak Lookout Tower.
Elev. 6600 ft.
Bottom of southwest-trending gully in large granite outcrop near ridgecrest (summit). Phenology: flowering.
with Patches
Accession: RM 698696, Barcode: RM0322955

Hackelia deflexa (1942)
BORAGINACEAE
Hackelia deflexa (Wahlenb.) Opiz
U.S.A., North Dakota, Ward County: Des Lacs National Wildlife Refuge: west side of Upper Des Lacs Lake, 8 mi N of Kenmare.
Weedy clearing. Phenology: flowering.
Accession: RM 604669

Hackelia deflexa var. americana (1988)
BORAGINACEAE
Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst.
U.S.A., Wyoming, Park County: Southwestern Absarokas: along Hardpan Creek, ca 24.5 air mi SW of Cody.
Streamside.
Accession: (unaccessioned)

Hackelia deflexa var. americana (2002)
Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst.
U.S.A., Wyoming, Campbell County: Thunder Basin National Grassland and Vicinity: ravine S of Bowman Hill on Rocky Point or County Road 85, ca 41 air mi NNE of Gillette; ca 24 air mi NE of Weston.
Sandy hills and ravines with ponderosa pine. Phenology: flowering & fruiting.
Peter J. Ebertowski, 1267. 7/7/2002.
Accession: RM (unaccessioned)

Hackelia deflexa var. americana (1991)
Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst.
U.S.A., Montana, Madison County: Madison Range: on the west front along Jack Creek Road, just W of bridge, ca 6 1/2 mi E of Ennis.
Elev. 5500 ft.
Shrubland adjacent to creek with Amelanchier utahensis, Shepherdia canadensis, Prunus virginiana, and Rhus trilobata. Phenology: flowering.
Accession: RM 738340

U.S.A., Montana, Madison County: Madison Range: Gallatin National Forest: Koch Basin on the east side of Koch Peak at head of Tumbledown Creek, ca 25 mi SE of Ennis.
Elev. 9200 ft.
Open, rocky whitebark pine woodland with Lupinus sericeus, Eriogonum umbellatum var. majus, and Potentilla fruticosa. Phenology: flowering.
Accession: RM 784037, Barcode: RM0057445

U.S.A., Colorado, Grand County: Rabbit Ears Range: Lindsey Creek to Grannys Nightcap, ca 18.5 air mi NNW of Kremmling and ca 23 air mi SE of Steamboat Springs.
Along creek. Phenology: fruiting.
Accession: RM (unaccessioned)

U.S.A., Idaho, Washington County: Payette National Forest and Vicinity: Cuddy Mountains Area: Hornet Reservoir, ca 15 air mi NW of Council; ca 16 air mi NNW of Cambridge.
Lakeside meadow. Phenology: flowering & fruiting.
with Denise L. Fairbanks
Accession: RM (unaccessioned)

U.S.A., Wyoming, Sheridan County: Wetlands and Adjacent Areas: Tongue River, ca 8 air mi SW of Ranchester.
Dry, rocky hillside. Phenology: flowering & fruiting.

U.S.A., Wyoming, Weston County: Wetlands and Adjacent Areas: seeping spring, ca 7 air mi NE of Newcastle.
Dry, sandy Pinus forest. Phenology: flowering & fruiting.

U.S.A., Wyoming, Sheridan County: Sheridan.
Wooded bottom lands. Phenology: fruiting.
Accession: RM 37559, Barcode: RM0323444

U.S.A., Wyoming, Washakie County: Red Bank.
43.6946° N, 107.2901° W; NAD 83, uncertainty 0.75 mi., Digital Map
Copses. Phenology: fruiting.

U.S.A., Wyoming, Campbell County:
Hills with pine, juniper, and Ribes. Phenology: flowering & fruiting.
Mary Alice Sanguinetti, 125. 7/7/1978.
with Keith H. Dueholm

U.S.A., Wyoming, Sheridan County: Hills E of Sheridan.
Moist shady draw. Phenology: flowering.

U.S.A., Wyoming, Weston County: Thompson Canyon.
Box elder with Potentilla and Chenopodium. Phenology: fruiting.

U.S.A., South Dakota, Meade County: Sturgis Tourist Park.
T5N R5E S10
Frequent in shade. Phenology: flowering.
Arthur C. McIntosh, 706. 7/8/1925.

U.S.A., South Dakota, Custer County: Jewel Cave National Monument: Lithograph Canyon between east boundary and Porthole [New Portal].
Roadside in canyon bottom, ponderosa pine forest. Phenology: flowering.
Hollis Marriott, 9733A. 7/4/1985.

U.S.A., South Dakota, Custer County: Jewel Cave National Monument: Prairie Dog Spring.
On mud below spring. Elev closer to 5200 f (USGS 7.5') G. Bryson Phenology: flowering.

U.S.A., South Dakota, Custer County: Jewel Cave National Monument: U.S. Hwy 16 just E of Hell Canyon.
Roadcut north of highway, among boulders. Phenology: flowering.

U.S.A., South Dakota, Harding County: Extreme northwest corner of county.
T23N R1E S36
Open woods. Phenology: flowering.
Annotations:
Annotated by C. L. Porter in 1949.
Accession: RM 69930

U.S.A., Colorado, Montezuma County: Mesa Verde National Park.
Among rocks. Phenology: fruiting.
Annotations:
1. Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst.
Annotated by B. E. Nelson.

U.S.A., Colorado, Ouray County: Ouray.
Phenology: flowering & fruiting.
Annotations:
1. Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst.
Annotated by Robert L. Carr.

U.S.A., Washington, Ferry County: Western Colville National Forest and Vicinity: Sherman Peak on Kettle Crest, ca 12 air mi ESE of Republic; Sherman Peak Loop Trail from Wash Hwy 20 at Sherman Pass.
Open slopes with granite outcrops, rocky ridge with talus; small lake with boggy area; mixed forest with subalpine fir, lodgepole pine, and larch. Phenology: flowering & fruiting.
Accession: (unaccessioned)

U.S.A., Washington, Ferry County: Western Colville National Forest and Vicinity: Kettle Crest Trail and Old Stage Road, ca 6 air mi N of Sherman Pass; from trailhead on Albion Hill Road.
South aspect sagebrush balds with scattered large Douglas fir; subalpine fir/lodgepole pine forest with large 1 year-old burn areas. Phenology: flowering & fruiting.
Accession: (unaccessioned)

U.S.A., Wyoming, Hot Springs County: Owl Creek/Bridger Mountains: Bridger Mountains: Klondike Peak and ridges NE and SW, NNW of Lysite Mountain; ca 24.5-25 air mi ESE of Thermopolis.
In limber pine forest on steep northerly facing slope.

U.S.A., Wyoming, Sheridan County: Powder River Basin: ca 6.5 air mi W of Parkman, ca 12 air mi NW of Dayton along East Pass Creek.
Roadside and riparian. Phenology: flowering & fruiting.

U.S.A., Wyoming, Washakie County: Big Horn Mountains: along Nowood River at the foot of Mahogany Butte, ca 27 air mi S of Ten Sleep, ca 9.5 air mi SSW of Big Trails.
Both entire and pinnatifid leaf forms together. Phenology: fruiting.

U.S.A., Wyoming, Campbell County: Powder River Basin: ca 6.5 air mi SW of Rockypoint.
Deep ravine in ponderosa pine covered uplift; ravine. Phenology: flowering & fruiting.

U.S.A., Wyoming, Sheridan County: [location and coordinates withheld]
Plains, flats along dry creekbed; near creekbed. Phenology: fruiting.

U.S.A., Wyoming, Sheridan County: Powder River Basin: ca 9 air mi WNW of Parkman.
Stream bottom, cherry thickets along hills; thickets. Phenology: fruiting.

U.S.A., Wyoming, Park County: Southwestern Absarokas: Ishawooa Creek, 2-4 mi W from trail head.
Douglas fir forests and stream bottoms with cottonwood.
with Neil Snow

U.S.A., Wyoming, Crook County: Black Hills: Bear Lodge Mountains: E of Norgrain Hill near South Dakota, ca 7.5 air mi NE of Aladdin; ca 26 air mi NE of Sundance.
In brush. Phenology: flowering & fruiting.

U.S.A., Wyoming, Crook County: Black Hills: Bear Lodge Mountains: E slope of Norgrain Hill near South Dakota, ca 7.5 air mi NE of Aladdin; ca 25.5 air mi NE of Sundance.
Moist shaded bottom of a draw.

U.S.A., Wyoming, Crook County: [location and coordinates withheld]
Oak woods, NE-facing slope. Phenology: flowering.

U.S.A., Wyoming, Crook County: [location and coordinates withheld]
Deciduous woods and thickets along creek. Phenology: flowering & fruiting.

U.S.A., Wyoming, Crook County: [location and coordinates withheld]
Oak woods at base of north-facing slope. Phenology: flowering & fruiting.

U.S.A., Wyoming, Crook County: Black Hills: Miller Creek, Garman Ranch; ca 9.5 air mi SSE of Devils Tower.
Deciduous woods along creek. Phenology: flowering & fruiting.

Deciduous thickets in north-trending draw. Phenology: flowering & fruiting.

U.S.A., Wyoming, Crook County: Black Hills: Goldie Divide near junction of Goldie and New Haven roads, ca 5 air mi NW of Hulett.
Deciduous woods in moist draw below reservoir. Phenology: fruiting.

U.S.A., Wyoming, Crook County: Devils Tower National Monument: southwest corner of monument.
On sandy soil at bottom of moist sandstone draw with Prunus virginiana and Acer negundo. Occasional; flowers white. Phenology: flowering.

U.S.A., Wyoming, Weston County: [location and coordinates withheld]
Deciduous thicket (plum-boxelder) in sagebrush-grassland (Artemisia cana). Phenology: flowering & fruiting.

Draw with oak, hawthorne, and wild plum. Phenology: flowering & fruiting.

U.S.A., Wyoming, Weston County: [location and coordinates withheld]
Aspen stand. Phenology: fruiting.

U.S.A., Wyoming, Weston County: Black Hills: Wet Parmlee Canyon near state line, ca 4.5 air mi SE of Four Corners.
Creek bottom in spruce-pine forest. Phenology: fruiting.
with Dennis Horning

U.S.A., Wyoming, Crook County: Black Hills: Devils Tower National Monument: trail E from Joyner Ridge parking lot.
Grassy ridge, sandstone outcrops and ponderosa pine understory. Phenology: fruiting.
with Hollis Marriot and Gael Fonken

U.S.A., Wyoming, Crook County: Black Hills: ca 3 air mi NNE of Aladdin.
Prairie, wooded stream bottom, and sandstone outcrops; wooded slope. Phenology: fruiting.

U.S.A., Montana, Gallatin County: Gallatin Range: Gallatin National Forest: just off Forest Road 62 by Window Rock Cabin, ca 15.5 air mi SSE of Bozeman.
Open meadow dominated by forbs and stream bank with beaver ponds. Phenology: fruiting.
with Ken Elliott
Accession: RM 868132, Barcode: USFS0062710

U.S.A., Montana, Meagher County: Lewis and Clark National Forest: Little Belt Mountains: South Fork Tenderfoot Road, ca 2 air mi from junction with Forest Road 589.
Elev. 1725 m.
46.9204° N, 111.0955° W; NAD 83, uncertainty 0.25 mi., GPS reading
Riparian zone and meadow NW of road; southwest aspect, 5-30% slope, limestone bedrock. Phenology: flowering & fruiting.
1. Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst..

U.S.A., Montana, Fergus County: Lewis and Clark National Forest: Big Snowy Mountains: Half Moon Canyon, within ca 1 air mi of Red Hill Road.
Limestone outcrops and coniferous forests. Phenology: fruiting.
1. Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst.
Annotated by Rick McNeill.

Annotated by Rick McNeill.

U.S.A., Montana, Park County: Absaroka Range: Gallatin National Forest: East Fork Boulder River valley, ca 41 air mi SSW of Big Timber; from the Box Canyon Trailhead over the Lake Plateau to the Upsidedown Creek Trailhead on Forest Road 6639.
Mixed conifer forest dominated by Pinus contorta and Pseudotsuga menziesii with a forb understory and with scattered meadows. Phenology: fruiting.
with Brian Elliott
Accession: RM 868133, Barcode: USFS0062711

U.S.A., Montana, Meagher County: Lewis and Clark National Forest: Little Belt Mountains: east side of Smith River, Indian Springs Campground, 26.8 air mi NW of White Sulphur Springs.
Elev. 1327 m.
Riparian zone and bench above river, wooded Pseudotsuga menziesii; west aspect; 0-5% slope. Phenology: flowering.
1. Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst..

U.S.A., Montana, Lewis and Clark County: Lewis and Clark National Forest: Front Ridge; ridge W of Green Fork Straight Creek, ca 1.5 air mi S of Cigarette Rock, ca 3/4 mi W of pond on Green Fork Straight Creek.
Elev. 2328 m.
Open rocky meadow, scattered Abies lasiocarpa and Pinus flexilis; southeast aspect; 0-10% slope; limestone. Phenology: fruiting.
1. Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I. M. Johnst..

U.S.A., Montana, Powder River County: Custer National Forest: Ashland District: vicinity of Cow Creek Camp/ Picnicground on Cow Creek ca 5.5 air mi W of Fort Howes Work Center, ca 25 air mi S Ashland.
Bushy valley bottom, stream bank, and base of ponderosa pine slope. Phenology: fruiting.
Accession: USFS 886442, Barcode: USFS0055145

U.S.A., South Dakota, Harding County: Custer National Forest: East Short Pine Hills: vicinity of Spring No. 3 just below and NE of Moreau Peak, ca 18 air mi SSW of Buffalo.
Narrow bushy draw with aspen, green ash, and chokecherry with wet areas. Phenology: fruiting.
Accession: USFS 886441, Barcode: USFS0055144

U.S.A., Montana, Carter County: Custer National Forest: Long Pines: at the mouth of Spring Canyon and just below at the northwest base of Rustler Ridge, ca 20.5 air mi SE of Ekalaka.
Bushy stream bank and adjacent moist bench in open ponderosa pine forest. Phenology: fruiting.
Accession: USFS 886440, Barcode: USFS0055143

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: just S of Forest Road 4785, ca 0.5 air mi SW of Camps Pass and ca 19 air mi ESE of Ashland.
Ponderosa pine and green ash-dominated draw. Phenology: fruiting.

U.S.A., Montana, Carter County: Custer National Forest: Long Pines: in and around Brewer Reservoir, just W of Forest Road 3036, ca 13.5 air mi WNW of Camp Crook, SD.
In and around small draw dominated by aspen and surrounded by open ponderosa pine woodland. Phenology: flowering & fruiting.
Accession: USFS (unaccessioned), Barcode: USFS0055975

U.S.A., South Dakota, Harding County: Custer National Forest: West Short Pine Hills: in and around generally west-facing draws ca 2.5 air mi SSW of JK Butte, ca 10 air mi SSE of Camp Crook.
Flowing water draw dominated by green ash and sedges. Phenology: flowering & fruiting.

U.S.A., South Dakota, Harding County: On the west side of The Jumpoff, ca 1 air mi E of County Road 897 and ca 3 air mi S of SD Hwy 20, ca 7.5 air mi ESE of Camp Crook.
Flat dominated by green ash. Phenology: fruiting.

U.S.A., South Dakota, Harding County: Custer National Forest: Slim Buttes: in and around draws SE of junction of North Divide or Forest Road 3125 and Forest Road 3145 in the vicinity of Camp Spring, ca 4.5 air mi S of Reva Gap, ca 19 air mi ESE of Buffalo.
In and near flowing water draw dominated by green ash, grasses, and sedges. Phenology: flowering & fruiting.

U.S.A., South Dakota, Harding County: Custer National Forest: Slim Buttes: just SW of Summit Pass on SD Hwy 79, ca 23.5 air mi SE of Buffalo.
Green ash-dominated flowing-water draw. Phenology: fruiting.

U.S.A., Montana, Carter County: Custer National Forest: Ekalaka Hills: in the vicinity of Russel Spring just S of Camp Needmore off of Forest Road 3104, ca 6.5 air mi SE of Ekalaka.
Mixed upland ponderosa pine, green ash, and aspen slope. Phenology: fruiting.

U.S.A., South Dakota, Harding County: Custer National Forest: South Cave Hills: in and around Timber Canyon, between Forest roads 3113 and 3137, ca 11.5 air mi N of Buffalo.
Green ash-dominated draw. Phenology: fruiting.

U.S.A., Montana, Carter County: Custer National Forest: Long Pines: just E of Forest Road 3117, ca 1/4 air mi SE of Iron Spring, ca 18 air mi NW of Camp Crook, SD.
Grass-dominated meadow into wind-thrown ponderosa pine woodland. Phenology: fruiting.
with G. Gust

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: N of Forest Road 4409, generally between Cabin Creek Spring and Deer Creek Spring, ca 11 air mi ENE of Ashland.
In and on margins of wet meadow dominated by sedges and grasses. Phenology: fruiting.

U.S.A., South Dakota, Harding County: Custer National Forest: Slim Buttes: in the vicinity of the head of Red Cross Draw, ca 1 air mi N of Reva Gap, ca 18 air mi E of Buffalo.
Moist hardwood-dominated draw. Sheet 2 of 2. Accession #886503 is sheet 1 of 2. Phenology: fruiting.

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: just N of Forest Road 4410, ca 2 air mi E of Yager Butte, ca 17 air mi ESE of Ashland.
In and around flowing-water hardwood draw with green ash, boxelder, chokecherry, snowberry, and Monarda. Phenology: fruiting.

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: in vicinity between Holiday Campground and SE to Sutton Spring, E of Forest Road 4777, ca 15 air mi ENE of Ashland.
In and around moist draw dominated by grasses and sedges. Phenology: fruiting.

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: in the vicinity of Griffin Pass, E of Forest Road 4431, ca 21.5 air mi ENE of Ashland.
Northeast-facing slope dominated by ponderosa pine and grasses with scattered rock outcrops. Phenology: fruiting.

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: in the vicinity of East Fork Spring, just S of East Fork Otter Creek Road, ca 14 air mi ENE of Ashland.
Moist, grazed draw dominated by green ash, Prunus, and Crataegus. Phenology: fruiting.

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: in the vicinity of Suicide Pass near Forest Road 4777, ca 1 air mi N of U.S. Hwy 212, ca 18.5 air mi E of Ashland.
Meadow dominated by grasses, snowberry, sagebrush, and Perideridia. Phenology: fruiting.

U.S.A., Montana, Phillips County: Ca 2-2.5 air mi E of Frenchman Creek and ca 0.5 mi S of Saskatchewan, ca 53 air mi NNE of Malta.
Prunus thicket on north-facing slope. Phenology: fruiting.

U.S.A., Montana, Phillips County: Upper end of Peck Coulee ca 4 air mi S of Saskatchewan and ca 3 air mi E of Frenchman Creek, ca 51 air mi NNE of Malta.
Buffaloberry thicket on east-facing slope near the bottom of the draw. Phenology: fruiting.

U.S.A., North Dakota, Ramsey County: Lisbon.
Phenology: flowering.

U.S.A., Montana, Big Horn County: CX Coal Site, along Squirrel Creek.
Riparian woodlands under treees in dense shade. Phenology: fruiting.
with L. M. Mayer

U.S.A., North Dakota, Barnes County: Valley City.
Phenology: flowering.

Hackelia deflexa var. americana ()
U.S.A., North Dakota: Devils Lake, Sweetwater Lake.
Near lake shore in sandy soil. Collection date not specified on label. Phenology: vegetative only.

U.S.A., Montana, Silver Bow County: Butte Highland Mountains.
T1N R8W S36
Around a rock outcropping on a dry knoll with prairie vegetation. Phenology: buds.
Klaus H. Lackschewitz, 9725. 7/22/1981.

U.S.A., North Dakota, Ramsey County: Devils Lake.
In open woods. Phenology: flowering.

U.S.A., North Dakota, Rolette County: St. Johns.
Open places in woods. Phenology: flowering.

U.S.A., North Dakota, Cass County: Kindred.
Woods. Phenology: flowering.
Wilhelm G. Solheim, 182. 7/4/1929.
with O. A. Stevens

U.S.A., Wyoming, Platte County: Wyoming Army National Guard Camp Guernsey Training Area: along South Guernsey Road, ca 1 road mi S of Trail Ruts Golf Course.
zone 13, 522671E, 4677171N; NAD 83, Specimen Label
High plains, at base of cliffs, in dense stand of trees and shrubs, with Populus deltoides, Acer negundo, Prunus virginiana, Fraxinus pensylvanica, Arctium minus, Aspergo procumbens. Urtica dioica, Galium aparine, and Nepeta cataria; alluvial deposits (Holocene); soils Mainter-Keeline fine sandy loams. USGS 7.5' Register Cliff Quad. Phenology: flowering.
with Donald L. Hazlett

U.S.A., Montana, Powder River County: Custer National Forest: Ashland Ranger District: just S of Forest Road 4785, ca 0.5 air mi SW of Camps Pass and ca 19 air mi ESE of Ashland.
Ponderosa pine and green ash-dominated draw. Phenology: fruiting.
Accession: (unaccessioned), Barcode: USFS0050208

U.S.A., South Dakota, Harding County: Custer National Forest: Slim Buttes: in the vicinity of the head of Red Cross Draw, ca 1 air mi N of Reva Gap, ca 18 air mi E of Buffalo.
Moist hardwood-dominated draw. Sheet 1 of 2. Accession #886504 is sheet 2 of 2. Phenology: fruiting.

U.S.A., Idaho, Custer County: Challis National Forest: Lost River Range: Methodist Creek narrows, 8.1 air mi ENE of Mackay.
Sandy or gravelly soil in mesic barren ground vegetation; 11° slope, E aspect; volcaniclastic geology. 26-75 individuals apparent. Phenology: flowering & fruiting.

U.S.A., Wyoming, Johnson County: Big Horn Mountains: North Fork Powder River Wilderness Study Area: east side of river, ca 27 mi NW of Kaycee.
Limestone side-canyon, bottoms; with Populus spp.

U.S.A., South Dakota, Pennington County: Black Hills: Smith Gulch, ca 1 mi SW of Rochford, ca 15 mi NW of Hill City.
T2N R3E S27 NE1/4
Forested margin of iron fen, in shade; with Equisetum arvense, Picea glauca, and Cornus canadensis. Uncommon.